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To identify the variables affecting haematological parameters and body mass, we used a backward stepwise selection procedure based on the restricted maximum likelihood ratio with successive removal of non-significant variables.

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Moult score was based on the last moulted primary feather score 1—

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Het Gelders Landschap en Kastelen, provided permission to collect the birds from their land.

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Studies on rats show that, in response to restraint, haematological parameters change after 6 minutes [ 66 ].

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Acknowledgments We would like to acknowledge Susanne Lubbe for her assistance with the behavioural observations. The time it took the bird to approach this perch and the shortest distance to the novel object within s were used to assess a score, ranging from 0 the bird did not land on the perch with the object to 5 the bird pecked the object. Although this suggests the mate plays an important role in female-female aggression, it is unknown how the presence of the mate affects the number of interactions between female great tits. View Article Google Scholar 3. Several studies have demonstrated a sex difference in response to stressors [ 18 — 22 ], with females showing a higher glucocorticoid secretion than males [ 21 ], which in turn can result in a higher heterophil count [ 6 ]. Some studies report that animals that lose fights subordinates show higher glucocorticoid levels than animals that win fights dominants [ 12 , 13 ], others demonstrate the contrary [ 14 , 15 ], whereas some find no difference [ 16 , 17 ].

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